In chimpanzees, personality and dominance often go hand in hand. The alphas aren’t always the biggest and strongest individuals; they’re often in charge because they want to be in charge. (Despite some of the benefits conferred, not everyone is interested in running for president.)
At CSNW, Annie is the lowest in the hierarchy. She also happens to be a little anxious. It’s impossible to know whether she’s anxious because she’s the lowest in the hierarchy, or whether she’s the lowest in the hierarchy because she’s an anxious individual. (The truth is probably some combination of those options.) Despite being the lowest in the hierarchy, Annie doesn’t have it too bad. She has to wait her turn at meals, and she defers to the more dominant chimps in passing. But she also gets away with a fair amount of sass – probably because the more dominant chimps aren’t threatened by her. She can take food straight out of Negra’s hands – assuming it’s not something Negra cared too much about – and she always lets the other chimps know when they’ve wronged her. As the years go by, we see less and less anxiety and more and more peace and contentment in her eyes.